Allan Wilson, a groundbreaking researcher and a lightning rod for controversy, revolutionized science and galvanized the scientific community through his quantitative biochemical approach to the history of evolution. Drawing upon the insights and recollections of those who knew Wilson best, this program—narrated by paleoanthropologist Tim White, codiscoverer of the hominid “Lucy”—correlates milestones of his remarkable career with his enduring contributions that range from molecular phylogenies of multiple species to an understanding of mechanisms underlying the mode and tempo of organismal evolution. Commentary by David Wake, professor emeritus of zoology at the University of California, Berkeley, and many others is featured. (41 minutes)